Right now, I can tell you that I have a message proving that directories will indeed get you penalized. I had a pr4 51 authority link on a directory site that was very popular. Google doesn't generally give you an hints on links, but they did and I have proof. They told me to take specific directory links down that we built to very relevant sections. Not only that, but even told me that my main url of my site I used as the keywords was still spam.
So, Google I feel is hand picking the directory sites like DMOZ, etc, etc. I also don't feel directory sites are that high of a quality in a link anyways. What can you really say in 30 or so words about your site.
I am not saying to stay away from them all, I am just saying be very careful, because I thought they were ok to use if you picked good ones too, but it's not.
Have a great night.
MB